These are due to the induction of specific lipolytic and fat-browning regulator genes, namely HSL, LPL, Agtl, UCP1, PRDM16 ... through the activation of the AMPK signalling pathway.
The process involved increase in the expression of lipolytic (HSL, LPL, and Agtl) and fat browning regulator genes (UCP1, PRDM16 ... protein expression via the AMPK signaling pathway.